2024-2025 Course Catalog

Please note: The information displayed here is current as of Saturday, November 23, 2024, but the official Course Catalog should be used for all official planning.

Biology

ProfessorsB. De Stasio (Dennis and Charlot Nelson Singleton Professor of Biological Sciences) (on leave term(s) III), E. De Stasio (The Raymond H. Herzog Professor of Science) (on leave term(s) III), J. Humphries (on leave term(s) I), J. Sedlock (on leave term(s) I, II, III)
Associate professorsK. Dickson (chair), A. Hakes, B. Piasecki, N. Wall
Visiting Assistant ProfessorsR. Jadin, K. Webb
LecturersU. Schultz, R. Theisen (Grant Evaluation Analyst Coord)

Students come to Lawrence with varied interests in the life sciences, thus course offerings in biology span the breadth of this exciting discipline and the biology major is flexible. Though each student may create a unique path to completing a major in biology, we encourage students to sample courses across the breadth of the field, including courses in ecology, organismal biology, and molecular genetics. The flexibility of the major allows students to include off-campus study into the undergraduate experience, whether that be the Lawrence University Marine Program or an experience that does not include the study of biology.

The department encourages an open-ended, original, experimental approach to life science. The research-rich approach begins in the introductory course sequence that students can enter with either BIOL 130: Cellular Form and Function or BIOL 135: Ecology, Evolution, and Society. In BIOL 130: Cellular Form and Function, in which all department faculty members participate, students design, conduct, and interpret their own research projects and present their results at a professional-style symposium at the end of the term. Both BIOL 135: Ecology, Evolution, and Society and BIOL 150: Organismal Form and Function, include three open-ended research modules in the laboratory portion of the course, and BIOL 135 also introduces students to field-based research. BIOL 280: Experimental Design and Statistics builds upon the research-rich introductory sequence and focuses on experimental design and statistical analysis of data. Students should plan to take BIOL 280 their sophomore year if possible.

Experimental work becomes progressively more sophisticated and creative in advanced courses. All courses are designed to develop students’ insights and capacity to synthesize information, and they include discussions, readings, field trips, lab work, and interactive class work in those areas most closely related to the competence of the faculty. Most courses feature intensive laboratory or field instruction in which students use advanced research equipment to explore modern biological concepts.

All biology faculty members conduct active research programs and employ students during the summer as research assistants, as well as supervise students undertaking independent research for credit during the academic year. Motivated students may approach faculty about laboratory or field research after their first or second year of study. Many students culminate their work in biology with significant original research. In recent years, several papers with students and faculty as co-authors have been published in professional journals. Topics have included aquatic food chain energetics, host-parasite and plant/animal interactions, gene expression, and molecular mechanisms of vertebrate development. Recent advances in biological research are presented in a series of talks (BIOL 600) by faculty and by scientists from other universities. All students complete a project (research, curriculum development, outreach, or a creative project) of their own design as part of our Senior Experience program and they present their work at our annual BioFest celebration of Senior Experience (see below for more details).

Students who have strong secondary interests in chemistry, geosciences, or physics may construct majors involving biology and one of the other three natural sciences, using the interdisciplinary major in the natural sciences or the biochemistry major.


Required for the major in biology

Students who major in biology will learn to explain both theory of and evidence for basic biological principles including energy metabolism, inheritance, evolution, physiology, and ecosystem structure and function. They will demonstrate critical analysis of primary research literature and conduct original research.

The major in biology requires:

  1. The following biology courses:
    1. BIOL 130: Cellular Form and Function
    2. BIOL 135: Ecology, Evolution, and Society
    3. BIOL 150: Organismal Form and Function
  2. BIOL 280: Experimental Design and Statistics
  3. CHEM 116: Principles of Chemistry: Energetics and Dynamics
  4. At least six six-unit courses in biology numbered 200 or above (excluding Senior Experience courses), of which at least four must be laboratory courses
  5. Completion of the biology Senior Experience, including:
    1. 2 units of BIOL 501: Professional Development in the Life Sciences
    2. 1 unit of BIOL 600: Recent Advances in Biology Seminar
    3. A student-directed project
    4. 5 units of BIOL 650: Biology Senior Capstone I
    5. 1 unit of BIOL 651: Biology Senior Capstone II

Note: Only two six-unit courses designated as tutorial, directed study, or independent study can be counted toward the major or minor requirements and only one of those can be counted toward the upper-level laboratory requirement. Online coursework cannot be transferred to earn biology credit.


Required for the interdisciplinary major in natural sciences in biology and physics or geosciences

Biology students who have strong secondary interests in physics or geosciences may construct a major involving biology and geosciences or physics using the interdisciplinary major in the natural sciences. Previous interdisciplinary combinations of biology and chemistry have been replaced by the biochemistry major.

The requirements for the interdisciplinary major with biology as the primary discipline are:

  1. The following biology courses:
    1. BIOL 130:  Cellular Form and Function
    2. BIOL 150:  Organismal Form and Function
    3. BIOL 280:  Experimental Design and Statistics
  2. The following physics courses:
    1. PHYS 141: Principles of Classical, Relativistic, and Quantum Mechanics and PHYS 151: Principles of Classical Physics; or
    2. PHYS 151: Principles of Classical Physics and PHYS 160: Principles of Modern Physics
  3. The following courses if geosciences is the secondary discipline:
    1. GEOL 110: Introductory Geology or GEOS 110: Introduction to the Geosciences
    2. GEOL/GEOS 210: History of the Earth and Life
  4. At least 10 six-unit courses in the natural sciences (biology, chemistry, geosciences, and physics) numbered 200 or above, with at least five in biology (of which at least three must be laboratory courses) and at least three in the secondary discipline
  5. Completion of the biology Senior Experience

Senior Experience in biology

Required: A student-designed project, 2 units of BIOL 501: Professional Development in the Life Sciences, 1 unit of BIOL 600: Recent Advances in Biology Seminar, 5 units of BIOL 650: Biology Senior Capstone I, 1 unit of BIOL 651: Biology Senior Capstone.

Purposeful advising in the spring of the sophomore year and attendance at the spring BioFest is meant to inspire sophomore students to think about what they might want to undertake as their culminating project in biology. During the Fall term of the junior year, students will take BIOL 501: Professional Development in the Life Sciences to prepare students for both the Senior Experience and life after Lawrence. Project work (research, internship, creation of a curricular module, draft of a grant proposal, draft of a children's book on biology, production of art about biology, or other creative project) generally will be undertaken prior to Winter term of the senior year and may be based on an internship, summer or academic year research, a tutorial, course work, independent study, or other student activities.

In the senior year, students will take 1 unit of BIOL 600: Recent Advances in Biology Seminar in the Fall term, 5 units of BIOL 650: Biology Senior Capstone I in the Winter term, and 1 unit of BIOL 651: Biology Senior Capstone II in Spring Term for BioFest. The purpose of the BIOL 650 course is to bring to culmination each student’s individual Senior Experience project and to place that project in an academic context. Each student prepares a paper that places their project into a biological context, compares it to our past and current understanding of the topic using primary literature, and summarizes the student’s project or results. Students will begin gathering and organizing academic resources for this paper in BIOL 600. In BIOL 650, students preparing a thesis for honors will prepare a significant portion of their thesis, while a student creating a visual product or curriculum will describe the biological underpinnings of the work and reflect on the production of the work itself, for example. The Senior Experience will culminate with a symposium, BioFest, during Spring Term, at which all students will present the results of their projects (or the project itself) as a poster, demonstration, or other visual display.


Required for the minor in biology

  1. The following biology courses:
    1. BIOL 130: Cellular Form and Function
    2. BIOL 135: Ecology, Evolution, and Society
    3. BIOL 150: Organismal Form and Function
  2. BIOL 280: Experimental Design and Statistics (students majoring in disciplines requiring a research methods and statistics course may request exemption from the BIOL 280 requirement).
  3. At least three six-unit courses in biology numbered 200 or above, of which at least two must be laboratory courses.

Note: Only two six-unit courses designated as tutorial, directed study, or independent study may be counted toward the major or minor requirements, and only one of these may be counted toward the upper-level laboratory requirement. On-line coursework cannot be transferred to earn biology credit.


Teacher certification in science (grades 4-12)

Students who want to teach life sciences at the secondary level should choose a broad range of biology courses that includes ecology, plant and animal organismal biology, and molecular and cellular biology, and should gain experience in both field and laboratory research. To receive a science license, a student must complete a major in one of the sciences (biology, chemistry, environmental science, geosciences, or physics) and at least one course in each of the other sciences. Students who plan to seek teacher certification should review the requirements in the Education section of the catalog and meet with the director of teacher education, preferably before the end of the sophomore year.


Advanced Placement

Students who have earned AP or IB credit in biology are exempt from BIOL 130: Cellular Form and Function and BIOL 131: Biology Learning Teams, but not the other introductory courses in the major. 

Students without AP/IB credit can opt to take a departmental exemption exam for either BIOL 130: Cellular Form and Function and/or BIOL 150: Organismal Form and Function if they feel well qualified based on previous coursework. Contact the biology department chair, Kim Dickson (kimberly.dickson@lawrence.edu), for more information.



Courses - Biology

BIOL 100: The Biology of Human Reproduction

An introductory course focusing on human reproduction to demonstrate some basic biological principles. The course includes discussion of cellular and organismal processes related to the development of human biological complexity. Current research in reproductive biology and its impact on the individual and society is considered. Lecture and laboratory. Primarily for non-science majors; credit not applicable to the biology major.
Units: 6
Also listed as Gender Studies 180

BIOL 103: Biotechnology and Society

An examination of basic biological principles underlying current biotechnology in the fields of human genetics and genetic engineering. Discussion of methods of basic scientific research, the impact of technology on society, and ethical problems in human and agricultural genetics. Credit not applicable to biology major. Weekly laboratories will introduce basic experimental methodology and procedures.
Units: 6

BIOL 130: Cellular Form and Function

An exploration of fundamental cellular processes in an evolutionary context including homeostasis, cell cycle, gene expression, energy transformation, inheritance, and multi-cellular development. Experimental approaches will be emphasized. Lecture and laboratory. BIOL 130 is primarily offered to serve students exploring majors in Biology, Biochemistry, Neuroscience, and Environmental Science. A student looking for a laboratory course in the natural sciences in order to fulfil a general education requirement, might be better served taking courses such as BIOL 103 or BIOL 100. PREREQUISITES: First-year or second-year standing: consent of instructor required for juniors and seniors. Simultaneous enrollment in BIOL 130 and BIOL 131 required for first- and second-year students.
Units: 6
Prerequisite: First-year or second-year standing: consent of instructor required for juniors and seniors. Simultaneous enrollment in BIOL 130 and BIOL 131 required for first- and second-year students.

BIOL 131: Biology Learning Teams

Students will engage in peer-led-team-learning in which small teams of students participate in a variety of learning, team-building activities, and problem-solving using current course content with upper-class facilitators. The course will include discussion of collaboration skills, intersectionality, and college-level learning. Simultaneous enrollment in BIOL 130 is required. BIOL 131 is required of all first- and second-year students in BIOL 130. Enrollment of students beyond their second year is optional. PREREQUISITES: Simultaneous enrollment in BIOL 130 BIOL 131 required for first- and second-year students
Units: 1
Prerequisite: Simultaneous enrollment in BIOL 130 BIOL 131 required for first- and second-year students

BIOL 135: Ecology, Evolution, and Society

This course introduces the mechanisms of biological evolution that have resulted in earth’s rich diversity of life. We will explore the ways in which populations change, species interact, communities are structured and ecosystems function through an evolutionary lens. Connections to global crises related to human-driven processes, such as climate change, land use change, and disease spread will make these basic concepts germane to our lives, and underscore the interdependence of human and environmental health. A weekly laboratory will provide opportunities to practice field ecology research and run evolutionary lab experiments. BIOL 135 is primarily offered to serve students exploring majors in Biology, Environmental Science, and Environmental Studies. A student looking for a laboratory course in the natural sciences in order to fulfil a general education requirement, might be better served taking courses such as BIOL 103 or BIOL 100.
Units: 6
Also listed as Environmental Science 135

BIOL 150: Organismal Form and Function

Development, morphology, physiology, and ecology of plants, animals, fungi and unicellular organisms will be compared in evolutionary context. Phylogenic relationships, ecological interactions, and ecosystem processes will be explored. Lecture and laboratory. PREREQUISITES: BIOL 130 or departmental examination
Units: 6
Prerequisite: BIOL 130 or departmental examination

BIOL 191: Directed Study in Biology

Directed study follows a syllabus set primarily by the instructor to meet the needs or interests of an individual student or small group of students. The main goal of directed study is knowledge or skill acquisition, not research or creative work. PREREQUISITES: To register student must complete the Student Initiated Course Form with consultation of supervising instructor.
Units: 1 TO 98
Prerequisite: To register student must complete the Student Initiated Course Form with consultation of supervising instructor.

BIOL 200: Animal Behavior

A lecture and field-study course examining the principles and problems of animal behavior. Subjects include orientation, feeding, locomotion, communication, escape in time and space, biological rhythms, mate choice, and aspects of social behavior, examined from evolutionary, ontogenetic, physiological, ecological, and ethological perspectives. Lecture and laboratory. PREREQUISITES: BIOL 150
Units: 6
Also listed as Environmental Studies 210
Prerequisite: BIOL 150

BIOL 211: Botany

An introduction to the study of plants with an emphasis on their structure, development, physiology, and diversity. Although this course is lecture-based, students will frequently interact with plants from the Lawrence University greenhouse.
Units: 6

BIOL 221: Entomology

Topics covered will include a survey of all of the clades of insects with information on the systematics, diversity, ecology, life history, behavior and unique characteristics of each lineage. Lecture material will be augmented with required field trips to collect local species (terrestrial and aquatic) and the creation of a personal collection of species following the format as is customary for museum collections. PREREQUISITES: BIOL 150 and sophomore standing
Units: 6
Prerequisite: BIOL 150 and sophomore standing

BIOL 225: Microbiology (Lecture Only)

A study of microbial life with an emphasis on prokaryotes. Microbial physiology is examined in the context of how unique characteristics allow microbes to exploit a vast diversity of environments, including the human body. Lecture Only. PREREQUISITES: BIOL 130 and BIOL 150, CHEM 116 recommended
Units: 6
Prerequisite: BIOL 130 and BIOL 150, CHEM 116 recommended

BIOL 226: Microbiology

A study of microbial life with an emphasis on prokaryotes. Microbial physiology is examined in the context of how unique characteristics allow microbes to exploit a vast diversity of environments, including the human body. Laboratory exercises introduce students to techniques used to safely study microorganisms. PREREQUISITES: BIOL 130 and BIOL 150, CHEM 116 recommended
Units: 6
Prerequisite: BIOL 130 and BIOL 150, CHEM 116 recommended

BIOL 229: General Ecology (Lecture Only)

An introduction to the interactions between organisms and the environment. Explores the role of physical, chemical and biotic processes--including human activities--in determining the structure and function of populations, communities, and ecosystems. Topics include resource availability, competition, predation, symbiosis and natural and anthropogenic disturbances such as disease, biological invasions, pollution and climate change. Lecture only.
Units: 6
Also listed as Environmental Studies 229

BIOL 230: General Ecology

An introduction to the interactions between organisms and the environment. Explores the role of physical, chemical and biotic processes--including human activities--in determining the structure and function of populations, communities, and ecosystems. Topics include resource availability, competition, predation, symbiosis and natural and anthropogenic disturbances such as disease, biological invasions, pollution and climate change. Lecture and laboratory.
Units: 6
Also listed as Environmental Studies 220

BIOL 235: Evolutionary Biology

A study of biological evolution, including natural selection, adaptation, the evolution of sex, speciation, extinction, and constraints on evolutionary change. Reading primary literature is emphasized. Two lectures and one discussion per week. PREREQUISITES: BIOL 130 or ANTH 141
Units: 6
Also listed as Environmental Studies 213
Prerequisite: BIOL 130 or ANTH 141

BIOL 240: Morphogenesis of the Vertebrates

An integrated lecture and laboratory course that undertakes the study of the structure and function of vertebrate organ systems through examination of morphology. Vertebrate ontogeny, phylogeny, and anatomy are addressed. PREREQUISITES: BIOL 150
Units: 6
Prerequisite: BIOL 150

BIOL 242: Comparative Physiology

A comparative study of the variety of solutions and adaptations diverse animals can make to similar problems — obtaining and transporting oxygen, maintaining water and salt balance, utilizing food, movement, and nervous and hormonal integration. Lecture and laboratory. PREREQUISITES: BIOL 150, CHEM 116 recommended
Units: 6
Prerequisite: BIOL 150, CHEM 116 recommended

BIOL 245: Conservation Biology

This course explores scientific concepts related to the conservation and restoration of Earth's biological diversity. Topics include patterns of species and ecosystem diversity, the relationship between biodiversity and ecosystem functioning, causes of extinction, assessing extinction risk, behavioral indicators, in-situ and ex-situ management strategies for endangered species, and ecosystem restoration. Lecture only. PREREQUISITES: BIOL 230 or BIOL 135
Units: 6
Also listed as Environmental Studies 245
Prerequisite: BIOL 230 or BIOL 135

BIOL 260: Genetics

A lecture and laboratory study of the principles of inheritance, gene expression, introductory genomics, sex determination, and the concepts of historical and modern eugenics and genetic engineering. PREREQUISITES: Sophomore standing and BIOL 130 or ANTH 141
Units: 6
Prerequisite: Sophomore standing and BIOL 130 or ANTH 141

BIOL 265: Biogeochemistry

This course explores fundamental cycles between earth's major reservoirs of nutrients such as nitrogen, phosphorus, carbon and water. Through lecture and group presentations, students will gain a solid understanding of the fundamentals of biogeochemical cycles and the mechanism underlying the biological transformations of those elements. Lecture and laboratory. PREREQUISITES: BIOL 130 or GEOL 110
Units: 6
Also listed as Environmental Studies 265, Geosciences 265
Prerequisite: BIOL 130 or GEOL 110

BIOL 275: Introduction to Biophysics

Biophysics explores processes at the interface between biology and physics. This course uses physical principles to build mathematical and computational models of living systems. Students will learn to create models, compare models with experimental data, and refine their understanding of the living systems using these physics-based models. PREREQUISITES: Physics 151 or BIOL 170/BIOL 280
Units: 6
Also listed as Physics 275
Prerequisite: Physics 151 or BIOL 170/BIOL 280

BIOL 280: Experimental Design and Statistics

An introduction to experimental and sampling design in the fields of biology and biochemistry, as well as methods of data analysis and interpretation. The connection between statistical analysis and experimental design will be emphasized. Topics include descriptive, exploratory, and confirmatory statistical analyses. Lecture and computer laboratory. This course was previously known as BIOL 170. PREREQUISITES: BIOL 150 or consent of instructor
Units: 6
Prerequisite: BIOL 150 or consent of instructor

BIOL 310: Human Anatomy

A course in human anatomy only intended for students entering the allied health professions (e.g. nursing, PA, PT, athletic training) or forensic anthropology. Students learn detailed anatomy using full-size human models. Students are expected to learn structures of the skeletal, muscular, nervous (sensory included), circulatory, digestive, respiratory, urogenital, and endocrine systems. Emphasis is on the anatomy, particularly in the laboratory component of the course, but basic physiology is also covered. PREREQUISITES: BIOL 150 or ANTH 141
Units: 6
Prerequisite: BIOL 150 or ANTH 141

BIOL 325: Cell Biology

Survey of the structure and function of eukaryotic cells, the basic functional unit of life. Correlation of cellular structures including organelles, proteins, and membranes with functions such as cellular communication, division, transport, movement, and secretory pathways will be analyzed. Lecture and laboratory. PREREQUISITES: BIOL 130 and BIOL 150, BIOL 170/BIOL 280 recommended
Units: 6
Prerequisite: BIOL 130 and BIOL 150, BIOL 170/BIOL 280 recommended

BIOL 330: Aquatic Ecology

The principles of the ecology of fresh waters, developed through discussions, laboratory, and field investigations of the functional relationships and productivity of biotic communities as they are affected by the dynamics of physical, chemical, and biotic parameters. Lecture and laboratory. PREREQUISITES: BIOL 150 or BIOL 170/BIOL 280 (or concurrent enrollment) or BIOL 230
Units: 6
Also listed as Environmental Studies 310
Prerequisite: BIOL 150 or BIOL 170/BIOL 280 (or concurrent enrollment) or BIOL 230

BIOL 335: Plant Ecology

This course emphasizes core concepts in ecology and evolution from the unique perspective of plants. Students will explore the interactions between plants and their environment over a range of scales; from individuals to populations and communities. Lecture and laboratory. PREREQUISITES: BIOL 170/BIOL 280
Units: 6
Also listed as Environmental Studies 340
Prerequisite: BIOL 170/BIOL 280

BIOL 340: Topics in Neuroscience

A study of the nervous system from the perspectives of psychology and biology. Topics vary year to year and may include glial cells, neural development, and the evolution of nervous systems and neurotransmitter systems. Lecture only. May be repeated when topic is different.

Topic for Fall 2024: Biopsychology of Parenting
PREREQUISITES: CHEM 116, BIOL 150 and one course in psychology; or PSYC 360 and one course in biology; or consent of instructor This discussion-based course will explore the biological and psychological phenomena behind differences in parental practices. Particularly, we will address the physiological, hormonal, ecological and social bases of these differences. We will consider a variety of animal species, including humans.

Units: 6
Also listed as Psychology 580, Neuroscience 580
Prerequisite: CHEM 116, BIOL 150 and one course in psychology; or PSYC 360 and one course in biology; or consent of instructor

BIOL 345: Terrestrial Wildlife Ecology

A hands-on course intended to demonstrate basic ecological principles using local terrestrial ecosystems. Field research projects will introduce students to methods in hypothesis development, experimental design, data collection, statistical analysis, and scientific writing and presentation. Research topics will include estimating population size, community structure, plant-animal interactions, and foraging behavior. Lecture and laboratory. PREREQUISITES: BIOL 150, BIOL 135, BIOL 280, and sophomore standing
Units: 6
Also listed as Environmental Studies 345
Prerequisite: BIOL 150, BIOL 135, BIOL 280, and sophomore standing

BIOL 354: Molecular Biology

An interdisciplinary examination of regulatory mechanisms leading to differential gene expression. Main topics include transcription, translation, gene and protein structure, and modern genomics. The application of current molecular techniques is emphasized throughout the course. Laboratory work is experimental in approach. Lecture and laboratory. PREREQUISITES: BIOL 130 and CHEM 115
Units: 6
Prerequisite: BIOL 130 and CHEM 115

BIOL 360: Introduction to Bioinformatics

An introduction to the methods and software used to analyze biological data. Through lecture and guided tutorials, students will learn the structure and organization of biological databases, explore methods for examining genomic and proteomic data sets, and examine specific case studies relating to evolution, drug discovery and human variation. PREREQUISITES: BIOL 260
Units: 6
Prerequisite: BIOL 260

BIOL 370: Human Physiology

An examination of how the body maintains homeostasis. The various physiological systems (e.g., respiratory and cardiovascular) will be studied at multiple levels of organization, from molecular and cellular to the macroscopic. This course is primarily aimed at students entering the health sciences. The course will have both lectures and a laboratory component. PREREQUISITES: BIOL 150
Units: 6
Prerequisite: BIOL 150

BIOL 375: Biostatistics

This analytical and writing course trains students on the use of advanced analytical methods common in biological data analysis. Students complete analyses in the R programming software and work on individual datasets. Extensive writing about analytical methods, appropriate applications of analyses, and interpretations of these analyses from a large portion of this course. Lecture only with a term-long project required. PREREQUISITES: BIOL 170/BIOL 280 or consent of instructor.
Units: 6
Prerequisite: BIOL 170/BIOL 280 or consent of instructor.

BIOL 380: Ecological Modeling

An integrated lecture and computer laboratory introduction to the process of developing mathematical descriptions of the interactions between components of a population, community, or ecosystem, and the use of computer simulation as a tool for understanding ecology and natural resource management. Topics include population growth, predator-prey and competitor interactions, biogeochemical cycling, and mass balance in ecosystems. PREREQUISITES: At least one of the following: BIOL 229, BIOL 230, BIOL 245, BIOL 330, BIOL 335 or BIOL 345
Units: 6
Also listed as Environmental Studies 380
Prerequisite: At least one of the following: BIOL 229, BIOL 230, BIOL 245, BIOL 330, BIOL 335 or BIOL 345

BIOL 385: Urban Ecology and Sustainability

This course explores the ecology happening all around us in urban spaces. Students will learn foundational ecological principles and observe how these impact natural resource management and sustainability in cities. Students will have multiple field-trips to observe ecology and sustainability initiatives in action around the Fox Cities. They will also work closely in reviewing sustainability planning and projects on campus and in the community. PREREQUISITES: BIOL135 or ENST 150
Units: 6
Also listed as Environmental Science 385
Prerequisite: BIOL135 or ENST 150

BIOL 390: Tutorial Studies in Biology

Individual investigations of problems in biology. PREREQUISITES: To register student must complete the Student Initiated Course Form with consultation of supervising instructor
Units: 1 TO 98
Prerequisite: To register student must complete the Student Initiated Course Form with consultation of supervising instructor

BIOL 391: Directed Study in Biology

Directed study follows a syllabus set primarily by the instructor to meet the needs or interests of an individual student or small group of students. The main goal of directed study is knowledge or skill acquisition, not research or creative work. PREREQUISITES: To register student must complete the Student Initiated Course Form with consultation of supervising instructor.
Units: 1 TO 98
Prerequisite: To register student must complete the Student Initiated Course Form with consultation of supervising instructor.

BIOL 395: Internship In Biology

The academic component of the internship includes readings related to the substance of the internship, discussions with the faculty supervisor, and a written report appropriate to the discipline. Course grades are based on this academic work.
Units: 1 TO 98

BIOL 399: Independent Study in Biology

Individual, in-depth investigation of a specific biological problem. Students contemplating an honors project should register for this course. PREREQUISITES: To register student must complete the Student Initiated Course Form with consultation of supervising instructor
Units: 1 TO 98
Prerequisite: To register student must complete the Student Initiated Course Form with consultation of supervising instructor

BIOL 414: Forest Ecology

This course will dive into the fundamentals of Forest Ecology, building off foundational knowledge in Earth and Environmental sciences. We will begin the course with discussions and intensive readings mixed in with primary scientific literature and talking with experts in forest ecology, including practitioners in the Department of Natural Resources and the Northeast Wisconsin and Door County land trusts. Students will build advanced practical skills of nature resource management in forest environments, while honing their speaking and written communication skills. A Bjorklunden trip to assess and design a forest ecological management plan is an essential part of this course (mandatory), will be complemented by field trips to local State Parks, Forests, and Conservation Organizations. Each student will present their final component of a forest ecology management plan, as a final term project. PREREQUISITES: PREREQUISITES: GEOS 110, GEOS 150 or BIOL 135 in addition to a 200-level course, or permission of instructor
Units: 6
Also listed as Geosciences 414, Environmental Science 414
Prerequisite: PREREQUISITES: GEOS 110, GEOS 150 or BIOL 135 in addition to a 200-level course, or permission of instructor

BIOL 420: The Geography of Life: Biodiversity in a Changing Planet

Earth is a dynamic and changing planet, comprised of tightly linked ecosystems and organisms. In this course we explore relationships between the biotic and abiotic drivers that influence the distribution of global diversity. We use large-scale datasets to develop practical skills for monitoring the responses of biodiversity to environmental change. PREREQUISITES: BIOL 150 and BIOL 170/BIOL 280; preferred but not required: BIOL 230 and BIOL 235
Units: 6
Also listed as Environmental Studies 420
Prerequisite: BIOL 150 and BIOL 170/BIOL 280; preferred but not required: BIOL 230 and BIOL 235

BIOL 430: Immunology

This course will cover the basic concepts of immunology, including differentiation of immune cells, antibody structure and function, antigen-antibody reactions, the major-histocompatibility complex, the complement system, immune responses to pathogens, allergies and auto-immune diseases, and comparative immunology. The course will also examine recent advances in the field through current peer-reviewed publications. The weekly laboratory will examine the basic questions, experimental subjects, and procedures of the field. PREREQUISITES: BIOL 130, BIOL 150, and junior standing; or consent of instructor
Units: 6
Prerequisite: BIOL 130, BIOL 150, and junior standing; or consent of instructor

BIOL 431: Immunology (lecture only)

This course will cover the basic concepts of immunology, including differentiation of immune cells, antibody structure and function, antigen-antibody reactions, the major-histocompatibility complex, the complement system, immune responses to pathogens, allergies and auto-immune diseases and comparative immunology. The course will also examine recent advances in the field through current peer-reviewed publications. Lecture only. PREREQUISITES: BIOL 130, BIOL 150, and junior standing; or consent of instructor
Units: 6
Prerequisite: BIOL 130, BIOL 150, and junior standing; or consent of instructor

BIOL 433: Ecological Energetics (Lecture Only)

Study of the transfer and transformation of energy or energy-containing materials between and within organisms and populations. Discussions will cover food web structure and function across marine, freshwater and terrestrial environments. Part of the Marine Biology Term. Lecture only. PREREQUISITES: BIOL 330, concurrent enrollment in BIOL 505 and 226 and consent of instructor
Units: 6
Also listed as Environmental Studies 411
Prerequisite: BIOL 330, concurrent enrollment in BIOL 505 and 226 and consent of instructor

BIOL 434: Ecological Energetics

Field and laboratory experimental investigations of the transfer and transformation of energy or energy-containing materials between and within organisms and populations of aquatic ecosystems. Part of the Marine Biology Term. Lecture and laboratory. PREREQUISITES: BIOL 330, concurrent enrollment in BIOL 505 and 226 and consent of instructor
Units: 6
Also listed as Environmental Studies 410
Prerequisite: BIOL 330, concurrent enrollment in BIOL 505 and 226 and consent of instructor

BIOL 444: Biochemistry I

An introduction to the study of biological processes at the molecular level with emphases on protein struction and function, enzyme mechanism and kinetics, fundamentals of physical biochemistry, and the chemistry of biological molecules, including carbohydrates, lipids, and nucleic acids. PREREQUISITES: CHEM 250 or concurrent enrollment, or consent of instructor
Units: 6
Also listed as Chemistry 340
Prerequisite: CHEM 250 or concurrent enrollment, or consent of instructor

BIOL 450: Special Topics in Biology

A course designed to offer students an opportunity to study important issues in biology not covered in other regularly offered classes. Activities may include reading and analysis of material from primary literature, consideration of interdisciplinary connections, and field and laboratory activities.




Topic for Fall 2024: Herpetology
PREREQUISITES: BIOL 130 and BIOL 150, or consent of instructor This course provides an in-depth exploration of the scientific study of amphibians and reptiles. Students will examine the global diversity of these fascinating creatures, delving into their natural history, ecology, evolution, and conservation. In addition to studying amphibians and reptiles, students will gain a broader understanding of various biological disciplines—including diversity, taxonomy, anatomy, physiology, and biogeography—through detailed examples involving these species. A portion of the course will be dedicated to understanding the species native to Wisconsin. By the end of the course, students will have a comprehensive understanding of the critical roles amphibians and reptiles play in their ecosystems and the challenges they face in the modern world.

Units: 6
Prerequisite: BIOL 130 and BIOL 150, or consent of instructor

BIOL 453: Developmental Biology

An experimental approach to animal development with laboratory and lecture emphasis on molecular and cellular processes of embryogenesis. Includes discussions of pattern formation, differentiation, cell interactions, gametogenesis and fertilization. Lecture and laboratory. PREREQUISITES: BIOL 150; and one of the following (or concurrent enrollment): BIOL 354, BIOL 444/CHEM 340, BIOL 260, or BIOL 325
Units: 6
Prerequisite: BIOL 150; and one of the following (or concurrent enrollment): BIOL 354, BIOL 444/CHEM 340, BIOL 260, or BIOL 325

BIOL 455: Biochemistry II

A continuation of Biochemistry I. A study of biological processes at the molecular level with an emphasis on metabolic pathways, recent advances in biochemical medicine, and biochemical aspects of gene replication, protein synthesis, molecular motors, and sensing. The course is divided between lecture and discussion and will rely heavily on current biochemical literature. PREREQUISITES: CHEM 340 or consent of instructor
Units: 6
Also listed as Chemistry 440
Prerequisite: CHEM 340 or consent of instructor

BIOL 465: Advanced Biotechnology

An advanced course that examines the ways in which fundamental principles of biochemistry and molecular biology are transformed into technologies that revolutionize basic science, industrial processes, medicine, and agriculture. Lectures, discussions and laboratory work will focus on current primary literature and novel research questions. PREREQUISITES: CHEM 340, BIOL 354, or consent of instructor
Units: 6
Prerequisite: CHEM 340, BIOL 354, or consent of instructor

BIOL 501: Professional Development in the Life Sciences

A course designed for juniors who plan to do the biology senior experience. This course focuses on career exploration, professional development, and preparation for senior experience. Weekly presentations will be given by faculty, staff, and visiting researchers. Requirements include a resume, cover letter, a mock interview, and an annotated bibliography. Students planning to study off-campus fall term of their junior year should contact their academic advisor. PREREQUISITES: Junior standing or consent of instructor
Units: 2
Prerequisite: Junior standing or consent of instructor

BIOL 505: Coral Reef Environments

Examines the ecology of coral reef environments. Lecture, laboratory, and field components. Part of the Marine Biology Term. Lecture and laboratory. PREREQUISITES: BIOL 330 and concurrent enrollment in BIOL 226 and BIOL 434
Units: 6
Also listed as Environmental Studies 505
Prerequisite: BIOL 330 and concurrent enrollment in BIOL 226 and BIOL 434

BIOL 570: Biological Physics

Develops and explores the physical principles underlying biological systems, with a particular emphasis on building quantitative models. Applies fundamental topics including thermodynamics, fluid dynamics, elasticity, and electrostatics to model molecular and cellular phenomena such as gene expression, virus assembly, DNA bending and stretching, and nerve impulses. PREREQUISITES: PHYS 151, and one of PHYS 230, CHEM 252, CHEM 340, or BIOL 354
Units: 6
Also listed as Physics 570
Prerequisite: PHYS 151, and one of PHYS 230, CHEM 252, CHEM 340, or BIOL 354

BIOL 590: Tutorial Studies in Biology

Individual investigations of problems in biology. PREREQUISITES: To register student must complete the Student Initiated Course Form with consultation of supervising instructor
Units: 1 TO 98
Prerequisite: To register student must complete the Student Initiated Course Form with consultation of supervising instructor

BIOL 591: Directed Study in Biology

Directed study follows a syllabus set primarily by the instructor to meet the needs or interests of an individual student or small group of students. The main goal of directed study is knowledge or skill acquisition, not research or creative work. PREREQUISITES: To register student must complete the Student Initiated Course Form with consultation of supervising instructor.
Units: 1 TO 98
Prerequisite: To register student must complete the Student Initiated Course Form with consultation of supervising instructor.

BIOL 599: Independent Study in Biology

Individual, in-depth investigation of a specific biological problem. Students contemplating an honors project should register for this course. PREREQUISITES: To register student must complete the Student Initiated Course Form with consultation of supervising instructor
Units: 1 TO 98
Prerequisite: To register student must complete the Student Initiated Course Form with consultation of supervising instructor

BIOL 600: Recent Advances in Biology Seminar

A multidisciplinary lecture series on modern biological theory and research. Students attend seminars and prepare short summaries or “reaction papers” on topics covered. Biology faculty members and visiting scientists in biological and allied fields present seminars relating their research to the broader aspects of their disciplines. Topics discussed within any academic year provide a comprehensive exposure to the current frontiers of biological research. May be repeated for a maximum of three units. PREREQUISITES: Senior standing and declared major in biology, or consent of instructor
Units: 1
Prerequisite: Senior standing and declared major in biology, or consent of instructor

BIOL 650: Biology Senior Capstone

Senior capstone in which students will benefit from direct input and feedback on their scientific writing and oral presentation skills as they complete senior experience projects and papers. Successful completion of BIOL 650 includes participation in BioFest, a symposium of biology senior experience projects during spring term. PREREQUISITES: Major in biology or biochemistry, or in neuroscience with departmental approval; and senior class standing or departmental approval
Units: 1 OR 5
Prerequisite: Major in biology or biochemistry, or in neuroscience with departmental approval; and senior class standing or departmental approval

BIOL 651: Biology Senior Capstone II

BIOL 651 is the culmination of the biology senior experience in which students prepare and present their final project (e.g., scientific poster, video, educational application) at Biofest. Students will receive explicit instruction on effective visual communication. Requirements include the completion of a biology senior experience product and participation in Biofest. PREREQUISITES: BIOL 650
Units: 1
Prerequisite: BIOL 650

BIOL 690: Tutorial Studies in Biology

Individual investigations of problems in biology. PREREQUISITES: To register student must complete the Student Initiated Course Form with consultation of supervising instructor
Units: 1 TO 98
Prerequisite: To register student must complete the Student Initiated Course Form with consultation of supervising instructor

BIOL 691: Directed Study in Biology

Directed study follows a syllabus set primarily by the instructor to meet the needs or interests of an individual student or small group of students. The main goal of directed study is knowledge or skill acquisition, not research or creative work. PREREQUISITES: To register student must complete the Student Initiated Course Form with consultation of supervising instructor.
Units: 1 TO 98
Prerequisite: To register student must complete the Student Initiated Course Form with consultation of supervising instructor.

BIOL 699: Independent Study in Biology

Individual, in-depth investigation of a specific biological problem. Students contemplating an honors project should register for this course. PREREQUISITES: To register student must complete the Student Initiated Course Form with consultation of supervising instructor
Units: 1 TO 98
Prerequisite: To register student must complete the Student Initiated Course Form with consultation of supervising instructor